if(!require(maxnet))
::install_github("BigelowLab/maxnet") devtools
Maxent SDM
SDM start to finish in one RMarkdown file
Set-up
You will need to install maxnet if you have not already. This checks if it is installed.
Load the necessary libraries. If you see errors that a library is not installed, you will need to install with install.packages("packagename")
.
suppressPackageStartupMessages({
library(maxnet)
library(dplyr)
library(maxnet)
library(sf)
library(stars)
library(geodata)
library(dismo)
library(lubridate)
library(sdmpredictors)
library(ggplot2)
library(cmocean)
library(janitor)
library(DT)
library(here)
library(rnaturalearth)
library(rnaturalearthdata)
library(raster)
library(ggspatial)
library(tidyverse)
library(robis)
})
Tell R that the root should be where this RMarkdown file resides. All our data files will be stored here.
<- "SDM"
sdm_dir ::i_am(paste0(sdm_dir,"/Turtle_maxnet.Rmd")) here
here() starts at /Users/eli.holmes/Documents/GitHub/tutorials_marine_sdm
Set up the spatial region
Create a bounding box
We create a bounding box using minimum and maximum coordinate pairs and assign a standared WGS 84 coordinate reference system. This creates a sfs_POLYGON.
<- sf::st_bbox(c(xmin = 41.875, xmax = 65.125,
extent_polygon ymax = -0.125, ymin = 32.125),
crs = sf::st_crs(4326)) %>%
::st_as_sfc() sf
Save the bounding box for future use.
<- here::here(sdm_dir, "sdm_data", "BoundingBox.shp")
fil ::write_sf(extent_polygon, fil) sf
Get the polygon in text format.
<- sf::st_as_text(extent_polygon[[1]])
pol_geometry pol_geometry
[1] "POLYGON ((41.875 32.125, 65.125 32.125, 65.125 -0.125, 41.875 -0.125, 41.875 32.125))"
Create a world map with our region
This allows us to check our polygon of interest is located in the correct region.
#Getting base map
<- rnaturalearth::ne_countries(scale = "medium", returnclass = "sf")
world
#Plotting map
<- ggplot() +
world_box #Adding base map
geom_sf(data = world) +
#Adding bounding box
geom_sf(data = extent_polygon, color = "red", fill = NA)+
#Setting theme of plots to not include a grey background
theme_bw()
world_box
Save the plot.
<- here::here(sdm_dir, "sdm_data", "world_box.rda")
fil save(world_box, file=fil)
Create a region map
Create a base map of our region and save it.
<- ggplot()+
base_region_map #Adding base layer (world map)
geom_sf(data = world, fill = "antiquewhite")+
#Constraining map to original bounding box
lims(x = c(st_bbox(extent_polygon)$xmin, sf::st_bbox(extent_polygon)$xmax),
y = c(sf::st_bbox(extent_polygon)$ymin, sf::st_bbox(extent_polygon)$ymax))
base_region_map
Save it
<- here::here(sdm_dir, "sdm_data", "base_region_map.rda")
fil save(base_region_map, file=fil)
We will add some more features to our map: colors, scale and compass.
<- base_region_map +
region_map #Add scale bar on the top right of the plot
annotation_scale(location = "tr", width_hint = 0.5)+
#Add north arrow on the top left of plot
annotation_north_arrow(location = "tl", which_north = "true",
#Include small buffer from plot edge
pad_x = unit(0.01, "in"), pad_y = unit(0.05, "in"),
#Set style of north arrow
style = north_arrow_fancy_orienteering) +
#Changing color, type and size of grid lines
theme(panel.grid.major = element_line(color = gray(.5), linetype = "dashed", size = 0.5),
#Change background of map
panel.background = element_rect(fill = "aliceblue")) +
labs(x = "longitude", y = "latitude")
Warning: The `size` argument of `element_line()` is deprecated as of ggplot2 3.4.0.
ℹ Please use the `linewidth` argument instead.
region_map
Scale on map varies by more than 10%, scale bar may be inaccurate
Save.
<- here::here(sdm_dir, "sdm_data", "region_map.rda")
fil save(region_map, file=fil)
We add some labels for the countries.
#Extracting labels for countries in base map
<- world %>%
world_points ::st_make_valid(world) %>%
sf#Getting centroids for all polygons in the world base map
::st_centroid(geometry) %>%
sf#Getting coordinates for each centroid
::st_coordinates() %>%
sf#Adding centroids to original base map
::bind_cols(world)
dplyr
#Do not use spherical geometry
::sf_use_s2(FALSE)
sf
#Adding labels to map
<- region_map +
region_map_label geom_text(data = world_points,
#Point to coordinates and column with country names
aes(x = X, y = Y, label = name),
#Changing color and size of labels
color = "darkblue", size = 3,
#Avoid label overlap
check_overlap = TRUE)
# Save
<- here::here(sdm_dir, "sdm_data", "region_map_label.rda")
fil save(region_map_label, file=fil)
#Checking final map
region_map_label
Loading in the saved files
Later when we need the extent polygon, we use
#Loading bounding box for the area of interest
<- here::here(sdm_dir, "sdm_data", "BoundingBox.shp")
fil <- sf::read_sf(fil) extent_polygon
We often will need a sf bbox (bounding box object). To create that use
<- sf::st_bbox(extent_polygon) bbox
We load the maps as
<- here::here(sdm_dir, "sdm_data", "region_map_label.rda")
fil load(fil)
Get occurrence data from robis
pol_geometry
is defined above.
pol_geometry
[1] "POLYGON ((41.875 32.125, 65.125 32.125, 65.125 -0.125, 41.875 -0.125, 41.875 32.125))"
Get the data. We use eval=redo
so that we do not redownload data if we do not need to.
<- FALSE redo
Set the species we want
<- c("Chelonia mydas", "Caretta caretta", "Eretmochelys imbricata", "Lepidochelys olivacea", "Natator depressus", "Dermochelys coriacea") spp
Download the data.
<- robis::occurrence(spp, startdate = as.Date("2000-01-01"), geometry = pol_geometry) obs
This has many columns that we don’t need. We reduce to fewer columns.
<- c("occurrenceID", "scientificName",
cols.to.use "dateIdentified", "eventDate",
"decimalLatitude", "decimalLongitude", "coordinateUncertaintyInMeters",
"individualCount","lifeStage", "sex",
"bathymetry", "shoredistance", "sst", "sss")
<- obs[,cols.to.use] obs
We also add a cleaner date with YYYY-MM-DD format.
$date <- as.Date(obs$eventDate) obs
Save our data.
<- here::here(sdm_dir, "sdm_data", "sdm_data_all.csv")
obs_csv ::write_csv(obs, obs_csv) readr
Clean and prep data
Clean and prepare the data for our model and save to a new file name.
Load data in
# presence data
<- here::here(sdm_dir, "sdm_data", "occ_all.csv")
fil <- read.csv(fil) occ_all
We will call the cleaned data occ
.
# subset the occurences to include just those in the water
<- occ_all %>%
occ subset(bathymetry > 0 &
> 0 &
shoredistance < 200)
coordinateUncertaintyInMeters
# seeing how often each species occurs
table(occ$scientificName)
Caretta caretta Chelonia mydas
5141 7060
After cleaning we discover that we only have loggerhead and green sea turtles. Also there are only juvenile loggerheads and we do not know the life-stage of the green turtles.
table(occ$lifeStage, occ$scientificName, useNA="ifany")
Caretta caretta Chelonia mydas
Juvenile 5141 0
<NA> 0 7060
Select columns and add a common name column.
colnames(occ)
[1] "occurrenceID" "scientificName"
[3] "dateIdentified" "eventDate"
[5] "decimalLatitude" "decimalLongitude"
[7] "coordinateUncertaintyInMeters" "individualCount"
[9] "lifeStage" "sex"
[11] "bathymetry" "shoredistance"
[13] "sst" "sss"
[15] "date"
We want these. The last two are sea surface temperature and salinity.
<- c("scientificName", "date", "decimalLatitude", "decimalLongitude", "lifeStage", "bathymetry", "sst", "sss") cols
Subset the columns.
<- occ %>% dplyr::select(all_of(cols)) occ.sub
Change the column names.
colnames(occ.sub) <- c("sci.name", "date", "lat", "lon", "life.stage", "bathy", "SST", "SSS")
Add common.name column.
<- occ.sub %>%
occ.sub mutate(common.name = case_when(sci.name == "Caretta caretta" ~ "Loggerhead",
== "Chelonia mydas" ~ "Green")) sci.name
Save the cleaned file
<- here::here(sdm_dir, "sdm_data", "occ_clean.csv")
fil if (redo) readr::write_csv(occ.sub, fil)
Create background data
We will get random samples from our region.
Get a marine raster layer
We just need one because we use this to sample lat/lons from the marine environment. sdmpredictors
will download many files so we need to specify a directory.
# set a default data directory
options(sdmpredictors_datadir = here::here(sdm_dir, "sdm_data"))
# choosing marine
<- sdmpredictors::list_datasets(terrestrial = FALSE, marine = TRUE)
env_datasets <- sdmpredictors::load_layers("MS_bathy_5m")
env_stack <- env_stack %>% raster::crop(extent_polygon) env_stack
Plot to check that the layer looks ok. This is bathymetry.
plot(env_stack)
Look at the raster to get some info on it.
env_stack
class : RasterBrick
dimensions : 388, 280, 108640, 1 (nrow, ncol, ncell, nlayers)
resolution : 0.08333333, 0.08333333 (x, y)
extent : 41.83333, 65.16667, -0.1666667, 32.16667 (xmin, xmax, ymin, ymax)
crs : +proj=longlat +datum=WGS84 +no_defs
source : memory
names : MS_bathy_5m
min values : -5468
max values : -1
Sample points from this
It returns a sf points object.
<- 1000
nsamp <- dismo::randomPoints(env_stack[[1]], nsamp)
absence colnames(absence) <- c("lon", "lat")
Make a plot.
<- absence %>%
absence_sf as_tibble() %>%
::st_as_sf(coords = c(x="lon", y="lat"), crs = 4326)
sf::mapview(absence_sf, col.regions = "gray") mapview
Warning in cbind(`Feature ID` = fid, mat): number of rows of result is not a
multiple of vector length (arg 1)
Save the absence locations to a file.
<- here::here(sdm_dir, "sdm_data", "absence.csv")
fil write.csv(absence, file=fil, row.names = FALSE)
Download sdmpredictors layers
Set datasets to marine.
<- sdmpredictors::list_datasets(terrestrial = FALSE, marine = TRUE)
datasets <- list_layers(datasets)
layers #View(layers) # if you want to view
Choose layers.
= c("BO_sstmean", "BO_bathymean", "BO22_ph", "BO2_dissoxmean_bdmean", "BO2_salinitymean_ss", "BO2_chlomean_ss", "BO21_nitratemean_ss") layercodes
Download layers.
<- sdmpredictors::load_layers(layercodes, rasterstack = TRUE)
env <- env %>% raster::crop(extent_polygon) env_crop
Look at our layers.
plot(env_crop)
Save the raster brick for later reloading.
<- stars::st_as_stars(env_crop) # convert to stars object
env.stars <- here::here(sdm_dir, "sdm_data", "env_stack.tif")
fil ::write_stars(env.stars, fil)
stars#y <- stars::read_stars(fil)
# We need to do this for sampling
<- terra::split(env.stars) env.stars
Environmental predictors for points
We will use the stars
package to sample from our raster layers.
Load in our point data as data frames.
# presence data
<- here::here(sdm_dir, "sdm_data", "occ_clean.csv")
fil <- read.csv(fil)
df.occ
# absence data
<- here::here(sdm_dir, "sdm_data", "absence.csv")
fil <- read.csv(fil) df.abs
Convert data frames to sf points objects. This is what stars
needs.
<- na.omit(df.abs) # just in case
df.abs <- sf::st_as_sf(df.abs, coords = c("lon", "lat"), crs = 4326)
sf.abs <- sf::st_as_sf(df.occ, coords = c("lon", "lat"), crs = 4326) sf.occ
Get environment values for the absence points. Each row in sf.abs
is a row in env.abs
.
<- stars::st_extract(env.stars, sf::st_coordinates(sf.abs)) %>%
env.abs ::as_tibble() %>%
dplyrna.omit()
head(env.abs)
# A tibble: 6 × 7
BO_sstmean BO_bathymean BO22_ph BO2_dissoxmean_bdmean BO2_salinitymean_ss
<dbl> <dbl> <dbl> <dbl> <dbl>
1 27.7 -4108 8.19 161. 36.0
2 29.6 -4243 8.19 176. 35.2
3 27.1 -3462 8.13 128. 36.5
4 26.1 -52 8.14 219. 38.9
5 26.4 -3037 8.14 135. 36.1
6 26.3 -856 8.13 4.43 36.2
# ℹ 2 more variables: BO2_chlomean_ss <dbl>, BO21_nitratemean_ss <dbl>
Get environment values for the occurence points. Each row in sf.occ
is a row in env.occ
.
<- stars::st_extract(env.stars, sf::st_coordinates(sf.occ)) %>%
env.occ ::as_tibble() %>%
dplyrna.omit()
head(env.occ)
# A tibble: 6 × 7
BO_sstmean BO_bathymean BO22_ph BO2_dissoxmean_bdmean BO2_salinitymean_ss
<dbl> <dbl> <dbl> <dbl> <dbl>
1 26.4 -59 8.17 183. 35.6
2 28.6 -3158 8.18 154. 35.7
3 26.4 -92 8.17 145. 35.6
4 26.8 -2764 8.15 130. 36.1
5 27.9 -4 8.13 198. 38.7
6 27.6 -8 8.13 198. 38.7
# ℹ 2 more variables: BO2_chlomean_ss <dbl>, BO21_nitratemean_ss <dbl>
Now make this into one data frame with a pa
column for 1 is a occurrence row and 0 if an absence row.
<- c(rep(1, nrow(env.occ)), rep(0, nrow(env.abs)))
pres <- data.frame(pa = pres, rbind(env.occ, env.abs))
sdm_data head(sdm_data)
pa BO_sstmean BO_bathymean BO22_ph BO2_dissoxmean_bdmean BO2_salinitymean_ss
1 1 26.431 -59 8.173 182.6581 35.63274
2 1 28.648 -3158 8.181 154.3245 35.73521
3 1 26.428 -92 8.173 145.4299 35.64719
4 1 26.822 -2764 8.148 130.0710 36.09258
5 1 27.851 -4 8.133 197.8163 38.67305
6 1 27.585 -8 8.133 197.6590 38.71139
BO2_chlomean_ss BO21_nitratemean_ss
1 0.337094 2.043796
2 0.107879 0.132562
3 0.343290 2.086616
4 0.217175 0.615217
5 0.140721 0.000003
6 0.141303 0.000003
Save to a file. We will use for other models.
<- here::here(sdm_dir, "sdm_data", "sdm_data.csv")
fil write.csv(sdm_data, row.names = FALSE, file=fil)
Fit Maxnet model
maxnet::maxnet(pres, environ)
pres
string of 1s and 0s for whether the row is a occurrence or a absence.environ
a data frame of the environmental variables only
<- sdm_data$pa
pres <- sdm_data %>% dplyr::select(-pa)
environ <- maxnet::maxnet(pres, environ) sdm.model
Model metrics
<- plot(sdm.model, type = "cloglog") responses
We have some bathymetry values > 0 which might be a problem.
table(environ$BO_bathymean>0)
FALSE TRUE
12617 277
Predicting
<- TRUE # see ?predict.maxnet for details
clamp <- "cloglog"
type <- sf::st_bbox(extent_polygon) # make a sf bounding box
bb <- predict(sdm.model,
predicted %>% sf::st_crop(bb),
env.stars clamp = clamp, type = type)
although coordinates are longitude/latitude, st_intersects assumes that they
are planar
predicted
stars object with 2 dimensions and 1 attribute
attribute(s):
Min. 1st Qu. Median Mean 3rd Qu. Max. NA's
pred 0.003818902 0.1137863 0.1655903 0.174387 0.2148277 0.9999996 52802
dimension(s):
from to offset delta refsys x/y
x 1 280 41.83 0.08333 +proj=longlat +datum=WGS8... [x]
y 1 388 32.17 -0.08333 +proj=longlat +datum=WGS8... [y]
Visualization
We can plot the predictions like so, but the default palette is not great and are map is distorted.
ggplot() +
geom_stars(data = predicted)
We can try the cmocean palette and fix the coordinates.
ggplot() +
geom_stars(data = predicted) +
scale_fill_cmocean() +
coord_equal()
Or other palettes and annotation.
<- ggplot() +
predplot geom_stars(data = predicted) +
scale_fill_cmocean(name = "ice", direction = -1, guide = guide_colorbar(barwidth = 1, barheight = 10, ticks = FALSE, nbin = 1000, frame.colour = "black"), limits = c(0, 1)) +
theme_linedraw() +
coord_equal() +
theme(panel.background = element_blank(),
panel.grid.major = element_blank(),
panel.grid.minor = element_blank()) +
labs(title = "Loggerhead and green sea turtle SDM in the Arabian Sea",
x = "Longitude",
y = "Latitude",
fill = "Probability",
shape = "Species (presence)",
subtitle = "Environmental predictors: mean SS temp, mean SS salinity, mean bathymetry, \nmean pH, mean DO, mean SS chlorophyll-a, mean SS nitrate")
predplot
# ggsave("SDM_loggerhead_green_w points.pdf", height = 6, width = 8.5)
Without the occurrence data points.
+
predplot geom_point(sf.occ, mapping = aes(shape = common.name, geometry = geometry), stat = "sf_coordinates", alpha = 0.3, color = "purple") +
geom_point(sf.abs, mapping = aes(geometry = geometry), stat = "sf_coordinates", alpha = 0.3, color = "green")
# ggsave("SDM_loggerhead_green.pdf", height = 6, width = 8.5)
Discussion
We did not do much cleaning of the data and we combined loggerheads and green sea turtles. We should separate these. Also some of the data are clearly tagging data and we should subsample that data to remove some of the temporal autocorrelation. We should also experiment with higher and lower numbers of background points.